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Nymphalidae - the brush-footed butterflies
Until the late 20th century the Nymphalidae, Libytheidae, Heliconiidae, Brassolidae, Amathusiidae, Morphidae, Ithomiidae, Acraeidae, Danaidae and Satyridae were regarded as true families and of equal status. The current view is that these should now be reduced in rank to become subfamilies, or in some cases mere tribes within the Nymphalidae.
Thus defined the Nymphalidae comprises about 6000 species divided into 12 sub-families, based on the most recent phylogenetic studies, published by Wahlberg in 2008. These subfamilies each have their own distinctive characteristics but are united by one common feature - their front pair of legs are greatly reduced in size and are useless for walking. In males they are reduced to such a degree that they are little more than hairy stumps, hence the popular term 'brush-footed butterflies'.
Wahlberg's classification is published in systematic order below :

LIBYTHEINAE

Libythea
Libytheana

DANAINAE

tribe

Danaini

Parantica
Miriamica

Ideopsis
Amauris
Tirumala
Danaus
Tiradelphe
Euploea
Idea
Protoploea
Lycorea
Anetia

 

tribe

Tellervini

Tellervo

tribe

Ithomiini

 

Elzunia
Tithorea
Aeria
Athesis
Eutresis

Athyrtis
Paititia
Olyras
Patricia
Melinaea
Methona
Thyridia
Scada
Sais
Forbestra
Mechanitis
Placidina
Aremfoxia
Epityches
Hyalyris
Napeogenes
Hypothyris
Pagyris
Ithomia
Megoleria
Hyposcada
Oleria
Ceratinia
Callithomia
Dircenna
Hyalenna

Episcada
Haenschia
Pteronymia
Velamysta
Godyris
Veladyris
Hypoleria
Brevioleria
Mcclungia
Greta
Heterosais
Pseudoscada

CHARAXINAE

tribe

Charaxini

Polyura
Charaxes

tribe

Euxanthini

Euxanthe

tribe

Pallini

Palla

tribe

Prothoini

Prothoe
Agatasa

tribe

Preponini

Agrias
Prepona
Archaeoprepona
Noreppa

tribe

Anaeini

Anaeomorpha
Coenophlebia
Zaretis
Siderone
Hypna
Anaea
Polygrapha
Consul
Memphis
Fountainea

MORPHINAE

tribe

Morphini

Morpho
Antirrhea
Caerois

tribe

Brassolini

Bia
Blepolenis
Brassolis
Caligo
Caligopsis
Catoblepia
Dasyophthalma

Dynastor
Eryphanis
Mielkella
Mimoblepia
Opoptera
Opsiphanes
Orobrassolis
Penetes
Selenophanes
Narope
Aponarope

tribe

Amathusiini

Faunis
Aemona
Melanocyma
Enispe
Discophora
Amathusia
Zeuxidia
Amathuxidia
Thaumantis
Thauria
Stichophthalma
Taenaris
Hyantis
Morphopsis

SATYRINAE

tribe

Elymniini

Elymnias
Elymniopsis

tribe

Haeterini

Cithaerias
Dulcedo
Haetera
Pierella
Pseudohaetera

tribe

Melanitini

Manataria
Cyllogenes
Melanitis
Parantirrhoea
Gnophodes

tribe

Dirini

Tarsocera
Dingana
Dira
Torynesis
Aeropetes
Paralethe

tribe

Zetherini

Callarge
Penthema
Zethera

Xanthotaenia
Ethope
Neorina

tribe

Satyrini

sub-tribes in bold font

Parargina

Chonala
Kirinia
Lasiommata
Lopinga
Orinoma
Pararge
Rhaphicera
Tatinga
Nosea

Lethina

Aphysoneura
Hanipha
Lethe

Enodia
Satyrodes
Neope
Ninguta
Ptychandra
Mandarinia
Pseudoneorina

Mycalesina

Admiratio
Bicyclus
Bletogona
Mycalesis
Nirvanopsis
Hallelesis
Henotesia
Heteropsis
Houlbertia
Masoura

Pseudomycalesis

Eritina

Coelites
Erites

Ragadina

Acrophtalmia
Acropolis
Ragadia

Coenonymphina

Coenonympha
Sinonympha
Lyela
Triphysa

Hypocystina

Erycinidia
Hyalodia
Altiapa
Harsiesis
Hypocysta
Lamprolenis
Platypthima
Paratisiphone
Argyronympha
Dodonidia
Argyrophenga
Percnodaimon
Erebiola
Argynnina
Geitoneura
Heteronympha
Nesoxenica
Oreixenica
Tisiphone
Zipaetis
Orsotriaena

Euptychiina

Amphidecta
Archeuptychia
Caenoptychia

Caeruleuptychia
Capronnieria
Cepheuptychia
Cercyeuptychia
Chloreuptychia
Cissia
Coeruleotaygetis
Cyllopsis
Erichthodes
Euptychia
Euptychoides
Forsterinaria
Guaianza
Godartiana
Harjesia
Hermeuptychia
Magneuptychia
Megeuptychia
Megisto
Moneuptychia
Neonympha
Oressinoma
Palaeonympha
Paramacera

Parataygetis
Pareuptychia
Paryphthimoides
Pharneuptychia
Pindis
Posttaygetis
Praefaunala
Pseudeuptychia
Pseudodebis
Rareuptychia
Satyrotaygetis
Splendeuptychia
Taydebis
Taygetina
Taygetis
Taygetomorpha
Yphthimoides
Zischkaia

Erebiina

Erebia

Maniolina

Aphantopus
Cercyonis
Hyponephele
Maniola
Proterebia
Pyronia

Melanargiina

Melanargia

Pronophilina

Altopedaliodes
Antopedaliodes
Apexacuta
Arhuaco
Calisto
Cheimas
Corades
Corderopedaliodes
Daedalma
Dangond
Drucina
Druphila
Eretris
Eteona
Foetterleia
Gyrocheilus
Ianussiusa
Junea

Lasiophila
Lymanopoda
Mygona
Neopedaliodes
Oxeoschistus
Panyapedaliodes
Paramo
Parapedaliodes
Pedaliodes
Pherepedaliodes
Physcopedaliodes
Praepedaliodes
Praepronophila
Proboscis
Pronophila
Protopedaliodes
Pseudomaniola
Punapedaliodes
Redonda
Sierrasteroma
Steremnia
Steroma
Steromapedaliodes
Thiemeia
Diaphanos
Idioneurula
Manerebia
Neomaniola
Sabatoga
Stuardosatyrus
Tamania
Argyrophorus
Auca
Chillanella
Cosmosatyrus
Elina
Etcheverrius
Faunula
Haywardella
Homoeonympha
Nelia
Neomaenas
Neosatyrus
Palmaris
Pampasatyrus
Pamperis
Punargenteus
Quilaphoetosus
Spinantenna
Tetraphlebia

Satyrina

Arethusana
Argestina
Aulocera
Berberia
Boeberia
Brintesia
Callerebia
Cassionympha
Chazara
Coenyra
Coenyropsis
Davidina
Hipparchia
Kanetisa
Karanasa
Loxerebia
Mashuna
Melampias
Minois
Neita
Neocoenyra
Neominois
Oeneis
Paralasa
Paroeneis
Physcaeneura
Pseudochazara
Pseudonympha
Satyrus
Strabena
Stygionympha
Ypthima
Austroypthima
Ypthimomorpha

CALINAGINAE

Calinaga

HELICONIINAE

tribe

Acraeini

Pardopsis
Acraea
Actinote
Altinote
Abanonote
Cethosia

tribe

Heliconiini

Philaethria
Podotricha
Dione
Agraulis
Dryadula
Dryas
Eueides
Heliconius

tribe

Argynnini

Euptoieta
Yramea

Childrena
Boloria
Issoria
Brenthis
Argynnis

tribe

Vagrantini

Terinos
Lachnoptera
Phalanta
Smerina
Vindula
Cirrochroa
Algiachroa
Algia
Cupha
Vagrans

LIMENITIDINAE

tribe

Limenitidini

Limenitis
Lelecella
Sumalia
Moduza
Patsuia
Lamasia
Parasarpa
Auzakia
Litinga
Pandita
Athyma
Adelpha
Tacoraea
Lebadea
Kumothales
Pseudacraea
Pseudathyma
Pseudoneptis

tribe

Neptini

Pantoporia
Lasippa
Neptis
Phaedyma
Aldania
Cymothoe
Harma

tribe

Parthenini

Parthenos
Bhagadatta

tribe

Adoliadini

Euthalia
Tanaecia
Neurosigma
Abrota
Dophla
Lexias
Bassarona
Catuna
Hamanumida
Aterica
Pseudargynnis
Cynandra
Euptera
Euryphura
Euryphaedra
Euphaedra
Euriphene
Bebearia

CYRESTINAE

tribe

Cyrestini

Cyrestis
Chersonesia
Marpesia

tribe

Pseudergolini

Pseudergolis
Dichorragia
Stibochiona
Amnosia

BIBLIDINAE

tribe

Biblidini

Biblis
Ariadne
Laringa
Eurytela
Neptidopsis
Mesoxantha
Byblia
Mestra
Archimestra
Vila

tribe

Epicaliini

Catonephele
Nessaea
Myscelia
Sea
Eunica
Sevenia
Cybdelis

tribe

Ageroniini

Hamadryas
Ectima
Panacea
Batesia

tribe

Epiphilini

Asterope
Pyrrhogyra
Epiphile
Lucinia
Bolboneura
Temenis
Nica
Peria

tribe

Eubagini

Dynamine

tribe

Callicorini

Diaethria
Callicore
Perisama
Orophila
Mesotaenia
Antigonis
Haematera
Paulogramma
Catacore

APATURINAE

tribe

Apaturini

Apatura
Mimathyma
Chitoria
Dilipa

Euapatura
Eulaceura
Rohana
Euripus
Helcyra
Herona
Hestina
Hestinalis
Sasakia
Sephisa
Apaturina
Apaturopsis
Timelaea
Thaleropis
Asterocampa
Doxocopa

NYMPHALINAE

tribe

Coeini

Historis
Baeotus
Pycina

tribe

Nymphalini

Colobura
Tigridia
Smyrna
Antanartia
Araschnia
Symbrenthia
Mynes
Hypanartia
Vanessa
Aglais
Nymphalis
Polygonia

tribe

Victoriniini

Anartia
Siproeta
Metamorpha
Napeocles
Rhinopalpa
Kallimoides
Vanessula

tribe

Junoniini

Junonia
Salamis
Yoma
Protogoniomorpha
Precis
Hypolimnas

tribe

Kallimini

Kallima
Doleschallia
Catacroptera
Mallika

tribe

Melitaeini

Euphydryas
Chlosyne
Microtia
Poladryas

Melitaea
Higginsius
Gnathotriche
Atlantea
Antillea
Phystis
Mazia
Ortilia
Phyciodes
Tegosa
Tisona
Anthanassa
Dagon
Telenassa
Eresia
Castilia
Janatella

 

 

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